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In this work, we study periodicity in strings with wildcards. A string T with at most k wildcards is called strongly (p,k)-periodic if the wildcards in T can be replaced with alphabet symbols to obtain a string with period p, and weakly (p,k)-periodic if T[i] matches T[i+p] for all i. Intuitively, both generalize to (≤ g, k)-periodicity, which is the property of being (p,k)-periodic for some p ∈ [1..g]. An ε-tester for a property 𝒫 is an algorithm that distinguishes between strings that satisfy 𝒫 and strings where one needs to change at least an ε-fraction of the symbols to obtain a string that satisfies 𝒫. We study one-sided error testers, where strings satisfying 𝒫 must always be accepted, while strings that are ε-far must be rejected with probability at least 2/3. The complexity of a tester is the worst-case number of symbols of an input of length n it must read to make the decision. We design the following testers for p,g ≤ n/2: 1) An ε-tester for strong (p,k)-periodicity with complexity Õ_ε(1) . 2) An ε-tester for strong (≤ g,k)-periodicity with complexity Õ_ε(√g). 3) An ε-tester for weak (p,k)-periodicity with complexity Õ_ε(min(k, n /(k+p))). 4) An ε-tester for weak (≤ g,k)-periodicity with complexity Õ_ε(min(k+ √{gk}, n/√k)). Additionally, we show a lower bound on the complexity of ε-testers for weak (≤ g,k)-periodicity, implying that our tester for weak (≤ g,k)-periodicity is optimal up to a multiplicative (ε^{-1} ln(gk))^O(1) factor for a wide range of g and k. Finally, our tester for strong (≤ g,k)-periodicity generalizes the one of [Lachish and Newman; Algorithmica 2011] for strings without wildcards, matching (up to polylogarithmic factors) the unconditional lower bound of ̃Ω(√g) in said work for constant ε.